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The Department of Biology offers a program leading to a Master’s degree in Science. Although there are no formal options, students are able to specialize in conservation biology, environmental microbiology, botany, cellular and molecular biology, ecology, physiology, genetics, mycology, microbiology (bacteriology), parasitology, virology and zoology.

In addition to the admission requirements of the Graduate Studies Office, a Bachelor of Science degree in Biology or its equivalent is required. Generally, only applicants with a minimum 3.00 GPA in biology courses will be considered for admission. Candidates must submit scores of the Graduate Record Examination (GRE), and the Test of English as a Foreign Language (TOEFL) scores (foreign applicants) as an additional requirement to be considered for admission.

Requirements for the Masters Degree in the Department of Biology are met with the approval of a minimum of thirty credit hours of graduate courses including the thesis, of which a maximum of nine can be advanced undergraduate (5000) courses. BIOL 6689 (Biological Research Methods) and BIOL 6690 (Graduate Seminar) are core courses required to all students. All other program requirements are those established by the Graduate Studies Office. All students are required to write a thesis, and to present a departmental seminar related to their thesis research prior to graduation.

Departmental facilities include laboratories dedicated to research in botany, cellular and molecular biology, comparative physiology, entomology, virology, microbiology, mycology, and other areas of biology; a herbarium and a greenhouse; zoological collections, and a laboratory of animal behavior. In addition, the Biology Department operates a Microscopy Center.

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ADVANCED UNDERGRADUATE COURSES


BIOL 5005. ELEMENTARY PLANT ANATOMY (II) (Even numbered years). Three credit hours. Two hours of lecture and one three-hour laboratory per week. Prerequisite: BIOL 3435 or BIOL 3417 or authorization of the Director of the Department.

The study of simple and complex tissues of the organs of vascular plants; the study of the characteristics of parenchyma, sclerenchyma and collenchyma cells, as well as the elements composing the xylem and phloem tissues.


BIOL 5007. GENERAL PLANT MORPHOLOGY (II) (Even numbered years). Three credit hours. Two hours of lecture and one three-hour laboratory per week. Prerequisite: BIOL 3417 or BIOL 3435 or authorization of the Director of the Department.

The general principles of plant morphology, including evolutionary tendencies, phylogenetic lines and the life cycles of the principal groups of plants.


BIOL 5009. PTERIDOLOGY (On demand). Three credit hours. Two hours of lecture and one three-hour laboratory per week. Prerequisite: BIOL 3417 or BIOL 3435 or authorization of the Director of the Department.

Lectures and laboratories on the morphology, taxonomy and ecological distribution of the local ferns and their allies. Assigned readings and field trips.


BIOL 5016. PLANT EVOLUTION (I) (Odd numbered years). Two credit hours. Two hours of lecture per week. Prerequisite: BIOL 3417 or BIOL 3435 or authorization of the Director of the Department.

Analysis of the geological, morphological, anatomical, physiological, and geographical evidence showing how the different plant phyla have evolved, with emphasis on the evolution of tracheophytes. Assigned reading reports.


BIOL 5017. TROPICAL BRYOLOGY (On demand). Three credit hours. Two hours of lecture and one three-hour laboratory per week. Prerequisite: BIOL 3417 or authorization of the Director of the Department.

The biology of mosses, liverworts, and hornworts, emphasizing the structure, identification, reproduction, and ecology of the native species of Puerto Rico. Field trips required.


BIOL 5018. PLANT PHYSIOLOGY (II). Four credit hours. Three hours of lecture and one laboratory of three hours per week. Prerequisites: BIOL 3417 or BIOL 3435 or authorization of the Director of the Department. Corequisite: QUIM 3032 or QUIM 3062 or QUIM 3463 or authorization of the Director of the Department.

Plant physiology: diffusion, transpiration, absorption and transport, mineral nutrition, metabolism, growth and development, hormones, effects of environmental factors.


BIOL 5045. SCANNING ELECTRON MICROSCOPY (SEM). Three credit hours. Two hours of lecture and one three-hour laboratory per week. Prerequisite: Authorization of the Director of the Department.

Theoretical and practical aspects of the scanning electron microscope (SEM) with emphasis on sample preparation for SEM, detection of the different types of signals emitted by the specimen, and image analysis.


BIOL 5397. EUKAKYOTIC MOLECULAR GENETICS. Four credit hours. Two hours of lecture and two four-hour laboratory per week. Prerequisites: (BIOL 3300 and QUIM 5071) or authorization of the Director of the Department.

Genome complexity; gene structure, regulation of transcription; mRNA processing; transposons; signal transduction; the genetics of development, the cell cycle, and cancer; research techniques in molecular genetics.


BIOL 5416. HERPETOLOGY (I). Three credit hours. Two hours of lecture and one three-hour laboratory per week.

A study of the biology, classification and morphology of amphibians and reptiles, with emphasis on local species. Field trips.


BIOL 5417. ICHTHYOLOGY (On demand). Three credit hours. Two hours of lecture and one three-hour laboratory per week.

A study of the biology, classification and morphology of fishes, with emphasis on local species. Field trips.


BIOL 5585. MEDICAL AND VETERINARY ENTOMOLOGY (I) (Even numbered years). Three credit hours. Two hours of lecture and one three-hour laboratory per week.

This course offers the student interested in entomology, animal husbandry or veterinary science, an opportunity to become familiar with the recognition, characteristics, habits and control of insects, ticks, mites, and other arthropods that attack man and domestic animals.


BIOL 5755. VIROLOGY (I). Three credit hours. Two hours of lecture and one three-hour laboratory per week. Prerequisite: BIOL 3770 or authorization of the Director of the Department.

The classification, structure, physiology and biochemical activities of viruses.


BIOL 5758. BACTERIAL GENETICS. Two credit hours. Two hours of lecture per week. Prerequisites: BIOL 3300 or BIOL 3770 or authorization of the Director of the Department.

DNA replication and expression in the prokaryotic cell; transfer of genetic information; the impact of genetic processes on the physiology and ecology of bacteria.


BIOL 5759. BACTERIAL GENETICS LABORATORY. Two credit hours. Two four-hour laboratory periods per week. Co-requisite: BIOL 5758.

Molecular techniques for the study of the genetics of bacteria and bacteriophages. Practical experiences in the processes of recombination, complementation, the control of genetic expression, and the transmission of genetic information among microorganisms.


BIOL 5765. MYCOLOGY (II) (Even numbered years). Three credit hours. Two hours of lecture and one three-hour laboratory per week. Prerequisite: BIOL 3770 or authorization of the Director of the Department.

A study of the morphology, physiology, classification and relation of fungi to man. Emphasis is given to the isolation and identification of the different groups.


BIOL 5815. ANIMAL BEHAVIOR (I). Three credit hours. Two hours of lecture and one three-hour laboratory per week.

A study of activities and responses of animals in meeting their life requirements. Field trips.


BIOL 5955. INTRODUCTION TO RESEARCH METHODS IN ECOLOGY (II) (Even numbered years). Three credit hours. One hour of lecture and two three-hour laboratory periods per week. Prerequisite: authorization of the Director of the Department.

Field and laboratory exercises serve to introduce the student to the basic methods used in ecological research. The student is trained in the use of computers for the analysis of ecological data.


BIOL 5990. FIELD BIOLOGY WORKSHOP (On demand). One to three credit hours. Thirty to sixty hours of workshop/practice per credit. Prerequisite: authorization of the Director of the Department.

Intensive practical experience in selected areas of field biology, in or outside of Puerto Rico. A final written report will be required.


Graduate Courses

BIOL 6015. INSECT MORPHOLOGY (I) (Odd numbered years). Four credit hours. Two hours of lecture and two two-hour laboratories per week.

A study of the general internal and external morphology of insects.


BIOL 6155. PLANT ECOLOGY (II) (Odd numbered years). Four credit hours. Two hours of lecture and two three-hour laboratories per week. Prerequisite: authorization of the Director of the Department.

The interrelations of plants and environment; climatic, edaphic, and biotic factors in their relation to origin, development, and structures of vegetation; introduction to ecological fieldwork and the methods of ecological research. Practice is given in the recognition of associations, determination, and description of their structure, and relationships and measurements of environmental factors. Reports required.


BIOL 6199. BEHAVIORAL ECOLOGY (II) (Odd numbered years). Three credit hours. Three hours of lecture per week.

Recent developments in behavioral ecology. Evolutionary and ecological models applied to the behavioral problems of survival and reproduction. Integration of theory with field and laboratory evidences. Field trips required.


BIOL 6356. CYTOGENETICS (II) (Even numbered years). Three credit hours. Two hours of lecture and one three-hour laboratory per week.

A study of different aspects of the cell that affect inheritance.


BIOL 6369. POPULATION GENETICS (II) (Even numbered years). Three credit hours. Three hours of lecture per week.

Genetic variation in natural populations of both plants and animals in different communities, covering selection, migration, mutations, mating systems, and the effect of population size on the maintenance of genetic variation.


BIOL 6605. ENVIRONMENTAL POLLUTION AND DISTURBANCE (II) (Even numbered years). Three credit hours. Two hours of lecture and one three-hour laboratory per week. Prerequisite: authorization of the Director of the Department.

An ecological consideration of pollution and disturbance of the environment; the effects of industrial, domestic and other pollutants of the ecosystem; the physical, chemical and biological parameters used in pollution control and abatement. Field trips.


BIOL 6607. POPULATION ECOLOGY (I) (Even numbered years). Three credit hours. Two hours of lecture and one three-hour laboratory per week.

Study of populations for analysis of the control and interaction among them. Topics such as mortality, fertility, population growth, competition and predator-prey interaction will be discussed.


BIOL 6610. LIMNOLOGY (II) (Odd numbered years). Three credit hours. Two hours of lecture and one three-hour laboratory per week.

A study of the physical, chemical and biological characteristics and interrelations of these factors in aquatic situations; community structure in still and running water; studies of local streams and ponds.


BIOL 6617. ADVANCED GENETICS (I) (Odd numbered years). Three credit hours. Two hours of lecture and one three-hour laboratory per week. Prerequisite: authorization of the Director of the Department.

Discussion of selected topics in genetics.


BIOL 6631. CELLULAR BIOCHEMISTRY AND PHYSIOLOGY (I). Four credit hours. Three hours of lecture and one three-hour laboratory per week.

The interconversions of energy in living cells; photosynthesis and carbohydrate metabolism as energy sources; the utilization of metabolic energy for protein synthesis; solute and solvent movements; nerve and muscle phenomena. Emphasis on metabolic regulation and enzyme action.


BIOL 6635. MEDICAL MYCOLOGY (I) (Odd numbered years). Three credit hours. Two hours of lecture and one three-hour laboratory per week. Prerequisite: authorization of the Director of the Department.

A study of the fungi pathogenic to man.


BIOL 6637. TAXONOMY AND MORPHOLOGY OF FUNGI (I) (Even numbered years). Three credit hours. Two hours of lecture and one three-hour laboratory per week.

A thorough coverage of the phycomycetes, ascomycetes, deuteromycetes and basidiomycetes from a taxonomical and morphological approach, with emphasis on saprophytes, zoopathogens and phytopathogens.


BIOL 6642. ADVANCED MYCOLOGY (I) (Odd numbered years). Three credit hours. Two hours of lecture and one three-hour laboratory per week.

A study of fungi with emphasis on current literature and methods of research.


BIOL 6650. BACTERIAL DIVERSITY. Three credit hours. Two hours of lecture and two hour-and-a-half laboratories per week.

The diversity of prokaryotic organisms in relation to ecophysiological and evolutionary perspectives, emphasizing their isolation, identification, and application.


BIOL 6688. SCIENTIFIC PHOTOGRAPHY FOR BIOLOGISTS (I). Two credit hours. Two three-hour laboratories per week. Prerequisite: authorization of the Director of the Department.

Photographic techniques in biological research. A presentation and a portfolio of the student's work are required.


BIOL 6689. BIOLOGICAL RESEARCH METHODS (I). Two credit hours. Two hours of lecture per week.

Methods and theory of investigation in the biological field, including study of the biological literature and sources of information from major institutions active in this field. The student is required to write a research proposal in his area of interest.


BIOL 6690. GRADUATE SEMINAR (I). One credit hour. Two hours of lecture per week.

Discussion of recent literature in biology and related fields. Students will discuss principal topics in their special fields.


BIOL 6705. ADVANCED FOOD MICROBIOLOGY (I) (Even numbered years). Three credit hours. Two hours of lecture and one three-hour laboratory per week.

Microbiology of food commodities. The nature and function of beneficial and harmful microorganisms. Foodborne diseases. Effects of food processing and storage on microorganisms.


BIOL 6990. RESEARCH (I, II). One to six credit hour periods per week.

Research for a thesis.


BIOL 6991-6992. SPECIAL STUDIES IN BIOLOGY (I, II)-(I, II). One to three credit hours per semester. One to three research periods per week each semester.

Supervised research in some special topics of biology other than a thesis problem, but designed to provide experience and training in scientific investigation.


BIOL 6993. SPECIAL TOPICS IN BIOLOGY I (On demand). One to six credit hours. One to six hours of lecture per week.

Selected topics in biology, botany, microbiology, and zoology.


BIOL 6994. SPECIAL TOPICS IN BIOLOGY II (On demand). One to six credit hours. One to six hours of lecture per week.

Selected topics in biology, botany, microbiology, and zoology.


BIOL 6997. SPECIAL TOPICS IN BIOLOGY: LABORATORY (On demand). One to six credit hours. One to six two-to four-hour laboratories per week.

Laboratory practice of selected topics in biology, botany, microbiology, and zoology.


Botany (BOTA)

BOTA 6006. PHYSIOLOGY OF BACTERIA (II). Three credit hours. Two hours of lecture and one three-hour laboratory per week.

The physiology of bacteria and the biochemistry of microbic processes.


BOTA 6007. PHYTOGEOGRAPHY (II) (Even numbered years). Three credit hours. Three hours of lecture per week. Prerequisite: authorization of the Director of the Department.

A study of the geographical distribution of plants, with special emphasis on the ecological and historical factors affecting their distribution.


Zoology (ZOOL)

Advanced Undergraduate Course

ZOOL 5005. INVERTEBRATES OF PUERTO RICO (I) (Odd numbered years). Three credit hours. Two hours of lecture and one-three hour laboratory per week.

Taxonomy and ecology of the most common invertebrates of Puerto Rico, especially Arthropoda (exclusive of insects and marine forms) and Mollusca. Field trips.


Graduate Courses

ZOOL 6019. ADVANCED PARASITOLOGY. Four credit hours. Three lectures and one three-hour laboratory per week. Prerequisite: authorization of the Director of the Department.

Lectures, conferences, reading and laboratory work dealing with practical problems of classification, morphology and host relations of animal parasites.


ZOOL 6025. SYSTEMATIC ZOOLOGY (II) (Even numbered years). Three credit hours. Three hours of lecture per week.

The naming and classification of animals, rules and basis of nomenclature, quantitative methods of analysis, and methods for presentation of systematic findings.


ZOOL 6039. ANIMAL ECOLOGY (On demand). Three credit hours. Two hours of lecture and one three-hour laboratory per week. Prerequisite: authorization of the Director of the Department.

A study of the principles of ecology as applied to animals.


ZOOL 6056. ZOOGEOGRAPHY (I) (Even numbered years). Three credit hours. Three hours of lecture per week.

A study of the geographical distribution of animals, with special emphasis on factors affecting this distribution. Assigned readings and reports.


ZOOL 6058. INSECT TAXONOMY (II) (Odd numbered years). Three credit hours. One hour of lecture and two two-hour laboratory periods per week.

The classification of the hexapoda. Construction of keys, preparation of description, nomenclatural problems, faunistic and monographic studies, catalogs and bibliographies.


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BIOLOGY FACULTY

A list of professors who engage in graduate activities in the Department follows including the highest earned degree, date, and institution granting the degree. Research and teaching interests are also included.

CARLOS ACEVEDO, Assistant Professor, Ph.D., 2006, Vanderbilt University School of Medicine. Research interests: Celular and Molecular Immunology. Teaching interests: Immunology, Celular and Molecular Biology.

DIMARIS ACOSTA-MERCADO, Assistant Professor, Ph.D., 2003, University of Guelph, Canada. Research interests: Ecology of Soil Protozoa, Biodiversity and Ecosystem Function. Teaching interests: Ecological and Research Methods, Protistology, Philosophy of Sciences, Microbial Ecology.

JAIME A. ACOSTA, Associate Professor, Ph.D., 1995, Virginia Polytechnic Institute and State University. Research interests: Entomology, Biocontrol of Hymenoptera, Ecology of Fire Ants. Teaching interests: Biology, Ecology, Zoology, Entomology.

MÓNICA ALFARO, Associate Professor, Ph.D., 2002, University of Puerto Rico. Research interests: Marine Biology, Ecology of Zooplankton. Teaching interests: Marine Ecosystems, Zoogeography, and Biological Sciences.

FERNANDO BIRD-PICÓ, Associate Professor, Ph.D., 1994, University of Kansas. Research interests: Herpetology, Ecology and Population Genetics, Conservation Biology. Teaching interests: Herpetology, Systematic Zoology, Population Genetics, Comparative Vertebrate Anatomy.

SONIA BORGES, Professor, D.Sc., 1988, Universidad Complutense de Madrid. Research interests: Systematics and Ecology of Terrestrial Oligochaetes (Annelida), Organic Waste Management by Vermicom-posting. Teaching interests: Zoology.

GARY J. BRECKON, Professor, Ph.D., 1974, University of California at Davis. Research interests: Taxonomy and Ecology of Antillian Plants, especially rare and endangered species. Teaching interests: Botany and Ecology.

LUCY BUNKLEY-WILLIAMS, Professor, Ph.D., 1984, Auburn University. Research interests: Fish Parasitology and Pathology, Parasite Taxonomy. Teaching interests: Parasitology, Zoology, Ichthyology.

ROSA BUXEDA, Professor, Ph.D., 1993, Rutgers University, New Brunswick. Research interests: Science Education, Microbial. Teaching interests: Microbial Physiology, Microbiology, Microbial Biochemistry.

MATÍAS CAFARO, Assistant Professor, Ph.D., 2003, University of Kansas, Lawrence. Research interests: Mycology, Symbiosis, interactions between arthropods and fungi. Teaching interests: Topics in symbiosis, Advanced Mycology, Systematics of Fungi, Molecular Systematics.

FRANKLIN CARRERO, Assistant Professor, Ph.D., 2005, University of Illinois, Urbana. Research interests: Cell and Developmental Neurobiology and Synaptogenesis. Teaching interests: Neurobiology, Cell and Developmental Biology, Cell Signaling and Communication.

MILDRED CHAPARRO, Professor, Ph.D., 1985, Texas A&M University. Research interests: Food Microbiology. Teaching interests: Microbiology, Food Microbiology.

JESÚS D. CHINEA, Associate Professor, Cornell University, Ph.D., 1992, Research interests: Ecology, Forest Dynamics, Exotic Species, Restoration Ecology, Applications of Remote Sensing and GIS to Ecology. Teaching interests: Ecology, Botany.

CARLOS A. DELANNOY, Professor, Ph.D., 1984, University of Colorado. Research interests: Endangered Species. Teaching interests: Ornithology, Ecology, and Conservation Biology.

NANETTE DIFFOOT-CARLO, Professor, Ph.D., 1992, Virginia Polytechnic Institute and State University. Research interest: Molecular Studies of Viral Replication. Teaching interests: Virology, Molecular Biology.

NICO M. FRANZ, Assistant Professor, Ph.D., 2005, Cornell University, Ithaca, New York. Research interest: Insect Systematics (Morphology, Phylogeny, Classification), Theory and Practice of Systematics, Philosophy of Sciences. Teaching interests: Insect Morphology, Insect Taxonomy, Advanced Methods in Biogeography and Systematics Insect Plant Interactions, Philosophy of Science.

DUANE A. KOLTERMAN, Professor, Ph.D., 1982, University of Wisconsin. Research interests: Conservation Biology, Plant Biosystematics, Plant Nomenclature. Teaching interests: Botany, Biological Nomenclature, Conservation Biology.

ALLEN R. LEWIS, Professor, Ph.D., 1979, University of Rochester. Research interests: Behavioral Ecology, Population Biology. Teaching interests: Ecology, Animal Behavior.

SANDRA L. MALDONADO-RAMÍREZ, Associate Professor, Ph.D., 2001, Cornell University, Ithaca, New York. Research interests: Mycology, Aerobiology, Plant Pathology, Fungal Endophytes. Teaching interests: Mycology.

JOSÉ A. MARI-MUTT, Professor, Ph.D., 1978, University of Illinois, Urbana. Research interests: Taxonomy of Apterygota. Teaching interests: Zoology, Entomology, Invertebrates.

JUAN C. MARTÍNEZ-CRUZADO, Professor, Ph.D., 1988, Harvard University. Research interests: Molecular Population Genetics and Evolution. Teaching interests: Genetics, Molecular Genetics.

ARTURO A. MASSOL, Professor, Ph.D., 1994, Michigan State University. Research interests: Biodegradation, Microbial Diversity, Environmental Microbiology. Teaching interests: Microbial Ecology, General Microbiology, Industrial Microbiology.

RAFAEL MONTALVO, Professor, Ph.D., 2003, University of Nebraska. Research interest: Extremophiles, Taxonomy, Physiology and Benetics of Archaca. Teaching interest: Micology.

VIVIAN NAVAS, Professor, Ph.D., 1990, University of Illinois, Urbana. Research interests: Cell and Structural Biology. Teaching interest: Biology, Histology, Electron Microscopy.

CARLOS A. PÉREZ-MUÑOZ, Professor, Ph.D., 1991, University of California, Davis. Research interests: Botany, Structural Biology, Plant Morphogenesis. Teaching interests: Biology, Botany, Plant Morphology, Plant Anatomy, Microtechniques, Scientific Photography for Biologists.

LUIS A. RÍOS-HERNÁNDEZ, Assistant Professor, Ph.D., 2003, University of Oklahoma. Research interests: Anaerobic processes, Syntrophic Metabolism. Teaching interests: General Microbiology, Microbial Physiology, Anaerobic Microbiology.

CARLOS RIOS-VELÁZQUEZ, Associate Professor, Ph.D., 2000, University of Wisconsin-Madison. Research interest: Bacterial Genetics and Physiology, Microbial Biotechnology and Bioprospecting. Teaching interest: Microbial Physiology and Genetics, Prokaryotic Molecular Genetics and Gene Regulation, Microbial Biotechnology.

JUAN A. RIVERO, Distinguished Professor, Ph.D., 1953, Harvard University. Research interests: Herpetology, Evolution and Ecology of Reptiles and Amphibians. Teaching interests: Herpetology, Evolution, and Sex Biology.

CARLOS M. RODRÍGUEZ-MINGUELA, Assistant Professor, Ph.D., 2005, Michigan State University. Research interest: Molecular Ecology of Antibiotic Resistance Genes, Development of Quantitiative Molecular Methods for Microbial Source Tracking, and the Ecology of Microbial Processes involved in the removal of Environmental Pollutants. Teaching interests: Microbiology, Microbial Ecology, Application of Bioinformatics to Molecular Microbial Ecology, Diversity of Prokaryotes.

ALEJANDRO RUIZ-ACEVEDO, Professor, Ph.D., 1981, University of Oklahoma. Research interests: Microbiology, Immunology, Medical Mycology. Teaching interests: Medical Mycology, Immunology, Clinical Microbiology, Industrial Microbiology.

CARLOS J. SANTOS-FLORES, Associate Professor, Ph.D., 2001, University of Wisconsin-Madison. Research interests: Limnology, Freshwater Ecology, Taxonomy of Algae and Microinvertebrates, Aquatic Fungi Taxonomy and Ecology. Teaching interests: Limnology, Freshwater Biology, Plankton Ecology.

INES SASTRE-DE JESÚS, Professor, Ph.D., 1987, City University of New York. Research interests: Systematics of Bryophytes, Bryophyte Ecology and Conservation. Teaching interests: Scientific Writing, Tropical Bryology, Plant Evolution, Botany.

DIMUTH SIRITUNGA, Assistant Professor, Ph.D., 2002, Ohio State University. Research interests: Plant Molecular Biology, Metabolic Engineering, Genetics. Teaching interest: Plant Molecular Biology, Plant Physiology, Genetics.

RICHARD D. SQUIRE, Professor, Ph.D., 1969, North Carolina State University. Research interests: Genetics and toxicology of the brine shrimp, Artemia. Teaching interests: Genetics, Citogenetics.

JOHN M. USCIAN, Professor, Ph.D., 1994, University of Nebraska. Research interests: Marine Fish, Biochemistry/Physiology. Teaching interests: Physiology, Cell Biology, Genetics.

MARÍA M. VARGAS, Professor, Ph.D., 1997, Arizona State University. Research interests: Mycology, Entomopathogenic fungi, Microscopy. Teaching interests: Mycology, Microbiology, Microscopy.

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